diff --git a/Cesium.Parser.Tests/PreprocessorTests/PreprocessorTests.cs b/Cesium.Parser.Tests/PreprocessorTests/PreprocessorTests.cs
index 8b2924be..71f859ce 100644
--- a/Cesium.Parser.Tests/PreprocessorTests/PreprocessorTests.cs
+++ b/Cesium.Parser.Tests/PreprocessorTests/PreprocessorTests.cs
@@ -909,13 +909,22 @@ public Task FileDefine() => DoTest(
     [InlineData("1 || 0", true)]
     [InlineData("1 || 1", true)]
 
-    // TODO[#532]: Need to add support for parsing negative numbers, now "-" is recognized as a separator
-    // [InlineData("-10 < 9", true)]
-    // [InlineData("-10 > 9", false)]
 
     [InlineData("0b11 == 3", true)]
     [InlineData("021 == 17", true)]
     [InlineData("0xF == 15", true)]
+
+
+    [InlineData("-10 > 9", false)]
+    [InlineData("-10 < 9", true)]
+    [InlineData("-10 < -9", true)]
+    [InlineData("-9 == -9", true)]
+    [InlineData("-9 <= -9", true)]
+    [InlineData("-9 >= 10", false)]
+    [InlineData(" -10 < 0x10", true)]
+    [InlineData(" -0 == 0", true)]
+    [InlineData(" -0 == +0", true)]
+    [InlineData(" -10 != +10", true)]
     public async Task EvaluateExpressionAllVariants(
         string expression,
         bool expectedResult)
diff --git a/Cesium.Preprocessor/CPreprocessorTokenType.cs b/Cesium.Preprocessor/CPreprocessorTokenType.cs
index b202ba4d..7a03802a 100644
--- a/Cesium.Preprocessor/CPreprocessorTokenType.cs
+++ b/Cesium.Preprocessor/CPreprocessorTokenType.cs
@@ -32,7 +32,7 @@ public enum CPreprocessorTokenType
     [Regex("[^ \t\v\f\r\n#;+\\-*/()=!<>\",.|\\[\\]&\\\\]+")]
     PreprocessingToken,
 
-    [Regex(@"([.]|[;+\\-*/=!,|&]+|<=|>=|>|<|[|])")]
+    [Regex(@"([.]|[;+\-*/=!,|&]+|<=|>=|>|<|[|])")]
     Separator,
 
     [Token("(")]
diff --git a/Cesium.Preprocessor/ConditionExpressions/BinaryExpression.cs b/Cesium.Preprocessor/ConditionExpressions/BinaryExpression.cs
index 8695049e..393dde47 100644
--- a/Cesium.Preprocessor/ConditionExpressions/BinaryExpression.cs
+++ b/Cesium.Preprocessor/ConditionExpressions/BinaryExpression.cs
@@ -48,7 +48,7 @@ public string EvaluateExpression(IMacroContext context)
 
         int Parse(Location location, string macrosValue)
         {
-            if (Regex.IsMatch(macrosValue, $"^(0|[1-9][0-9]*)$"))
+            if (Regex.IsMatch(macrosValue, $"^(-?|\\+?)(0|[1-9][0-9]*)$"))
                 return int.Parse(macrosValue);
 
             if (Regex.IsMatch(macrosValue, "^0b[01]+$"))
diff --git a/Cesium.Preprocessor/ConditionExpressions/UnaryExpression.cs b/Cesium.Preprocessor/ConditionExpressions/UnaryExpression.cs
index f512c935..26e66ccf 100644
--- a/Cesium.Preprocessor/ConditionExpressions/UnaryExpression.cs
+++ b/Cesium.Preprocessor/ConditionExpressions/UnaryExpression.cs
@@ -19,6 +19,8 @@ public string EvaluateExpression(IMacroContext context)
         return Operator switch
         {
             CPreprocessorOperator.Negation => !expressionValue.AsBoolean(Location) ? "1" : "0",
+            CPreprocessorOperator.Sub => $"-{expressionValue}",
+            CPreprocessorOperator.Add => $"{expressionValue}",
             _ => throw new CompilationException($"Operator {Operator} cannot be used in the preprocessor directives")
         };
     }