diff --git a/src/ocaml_compiler.ml b/src/ocaml_compiler.ml
index e6d5becf..b05fe920 100644
--- a/src/ocaml_compiler.ml
+++ b/src/ocaml_compiler.ml
@@ -30,6 +30,11 @@ let forpack_flags arg tags =
     Ocaml_arch.forpack_flags_of_pathname arg
   else N
 
+let include_flags_for_deps deps =
+  let dirnames = List.union [] (List.map Pathname.dirname deps) in
+  let include_flags = List.fold_right ocaml_add_include_flag dirnames [] in
+  S include_flags
+
 let ocamlc_c tags arg out =
   let tags = tags++"ocaml"++"byte" in
   Cmd (S [!Options.ocamlc; A"-c"; T(tags++"compile");
@@ -37,7 +42,7 @@ let ocamlc_c tags arg out =
 
 let ocamlc_link flag tags deps out =
   Cmd (S [!Options.ocamlc; flag; T tags;
-          atomize_paths deps; A"-o"; Px out])
+          include_flags_for_deps deps; atomize_paths deps; A"-o"; Px out])
 
 let ocamlc_link_lib = ocamlc_link (A"-a")
 let ocamlc_link_prog = ocamlc_link N
@@ -70,19 +75,17 @@ let ocamlopt_c tags arg out =
 
 let ocamlopt_link flag tags deps out =
   Cmd (S [!Options.ocamlopt; flag; forpack_flags out tags; T tags;
-          atomize_paths deps; A"-o"; Px out])
+          include_flags_for_deps deps; atomize_paths deps; A"-o"; Px out])
 
 let ocamlopt_link_lib = ocamlopt_link (A"-a")
 let ocamlopt_link_shared_lib = ocamlopt_link (A"-shared")
 let ocamlopt_link_prog = ocamlopt_link N
 
 let ocamlopt_p tags deps out =
-  let dirnames = List.union [] (List.map Pathname.dirname deps) in
-  let include_flags = List.fold_right ocaml_add_include_flag dirnames [] in
   let mli = Pathname.update_extensions "mli" out in
   let cmd =
     S [!Options.ocamlopt; A"-pack"; forpack_flags out tags; T tags;
-       S include_flags; atomize_paths deps;
+       include_flags_for_deps deps; atomize_paths deps;
        A"-o"; Px out] in
   if (*FIXME true ||*) Pathname.exists mli then Cmd cmd
   else
diff --git a/src/ocaml_specific.ml b/src/ocaml_specific.ml
index 35f0f3d9..79b37a40 100644
--- a/src/ocaml_specific.ml
+++ b/src/ocaml_specific.ml
@@ -393,12 +393,12 @@ rule "ocaml: cmx & o -> cmxa & a"
 
 rule "ocaml: p.cmxa & p.a -> p.cmxs & p.so"
   ~prods:["%.p.cmxs"; x_p_dll]
-  ~deps:["%.p.cmxa"; x_p_a]
+  ~deps:["%.p.cmxa"]
   (Ocaml_compiler.native_shared_library_link ~tags:["profile";"linkall"] "%.p.cmxa" "%.p.cmxs");;
 
-rule "ocaml: cmxa & a -> cmxs & so"
+rule "ocaml: cmxa -> cmxs & so"
   ~prods:["%.cmxs"; x_dll]
-  ~deps:["%.cmxa"; x_a]
+  ~deps:["%.cmxa"]
   ~doc:"This rule allows to build a .cmxs from a .cmxa, to avoid having \
         to duplicate a .mllib file into a .mldylib."
   (Ocaml_compiler.native_shared_library_link ~tags:["linkall"] "%.cmxa" "%.cmxs");;
diff --git a/testsuite/internal.ml b/testsuite/internal.ml
index 8a13b55d..5198579b 100644
--- a/testsuite/internal.ml
+++ b/testsuite/internal.ml
@@ -167,6 +167,32 @@ let () = test "OutputShared"
          T.f "_tags" ~content:"<*.so>: runtime_variant(_pic)"]
   ~targets:("hello.byte.so",["hello.native.so"]) ();;
 
+let () = test "CmxsStubLink"
+  ~options:[`no_ocamlfind]
+  ~description:".cmxs link rules pass correct -I flags"
+  ~tree:[T.d "src" [
+           T.f "foo_stubs.c" ~content:"";
+           T.f "libfoo_stubs.clib" ~content:"foo_stubs.o";
+           T.f "foo.ml" ~content:"";
+         ];
+         T.f "_tags" ~content:"
+<src/foo.{cma,cmxa}> : record_foo_stubs
+<src/foo.cmxs> : link_foo_stubs";
+         T.f "myocamlbuild.ml" ~content:"
+open Ocamlbuild_plugin
+let () =
+  dispatch begin function
+  | After_rules ->
+      dep [\"record_foo_stubs\"] [\"src/libfoo_stubs.a\"];
+      flag_and_dep
+        [\"link\"; \"ocaml\"; \"link_foo_stubs\"] (P \"src/libfoo_stubs.a\");
+      flag [\"library\"; \"ocaml\";           \"record_foo_stubs\"]
+        (S ([A \"-cclib\"; A \"-lfoo_stubs\"]));
+  | _ -> ()
+  end
+"]
+  ~targets:("src/foo.cmxs",[]) ();;
+
 let () = test "StrictSequenceFlag"
   ~options:[`no_ocamlfind; `quiet]
   ~description:"strict_sequence tag"