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Otherwise, size the item into the available space using its used flex basis in place of its main size, treating a value of content as max-content.
The phrase "in place of its main size" is confusing and should be removed.
The used flex basis is always either the main size (when flex-basis is initially auto) or a user-declared size that overrides the main size. Therefore, used flex basis is not an alternative to the main size—it's the actual size under consideration.